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activity adolescent female adult females adult males aggressive competition agonistic agonistic behavior alpha male animals apes association attack baboons behavior birth Bygott calls charging displays Chausiku chimpan consort consortship copulation dominance encounter estrous cycles estrous females Faben feeding station Figan Figure followed foraging frequently fruit Gombe Gombe National Park Goodall gorillas home range human immature individuals infant interac interactions involved Jane Goodall Jomeo juvenile K-group Kahama males Kajabala Kamemanfu Kasakela Kasakela males Kasanga Kasonta male's mating mature males monkeys months mother mountain gorilla moved neighboring nest Nishida nulliparous observed occurred offspring orangutans pant-hoots party patrols patterns period play pongids population primates pygmy chimpanzees rank recorded relationships reproductive resident male riverine forest Schaller seems seen sexual sexual dimorphism siblings silverback social grooming social structure species study area subadult males subgroups Tanjung Puting Tanzania termites tion tree Ugalla area unhabituated unit-group vocal Wrangham young