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activity adapted adult aggressive animals apes apparently baboons baby becomes begin behavior birds branch cage calls child chimpanzees close communication complex Courtesy depends dominance early ecology environment experiments face fact feeding females Figure forest fruit gestures gibbon give gorilla Grade grooming ground guenon hand howler human individuals infant language langur learning least leaves lemur less live macaques males mammals manipulation mating means measured monkeys months mother move natural nest normal objects observed original particularly pattern period play points predators primates probably problem range reach relative response rhesus savanna season seems separation sexual similar single situation social space species squirrel stage status sticks structure symbols Table territory tests theory threat trees troop turn usually whereas whole wild World World monkeys young