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activity acts adult females adult males aggression animals appear approach baboons begins behavior birth body cage central changes characteristics chimpanzees communication continue cycle described disease displays dominance effect example experience extremely feeding females field forest four frequently fruit function gorilla grooming habitat home range howlers human important increase individual infant infection interactions involved island Japanese leader learning least leaves less limited living macaques maintain major mating meters monkeys months mother move movements natural observer occur organization Panama particularly pattern percent performed perhaps period plants play population predation primates produce records relationships relatively response result role season seems separate sexual signals situation sleeping social sound species spider status structure tion trees troop units usually vervets vocalizations World World monkeys young