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allele frequencies allozyme amino acid assortative mating assume assumptions average calculate Chapter chromosome common consider decrease deleterious mutations disassortative mating disease allele DNA sequences dominance Drosophila melanogaster equilibrium estimate evolution evolutionary example expected heterozygosity favored females Figure gamete gametic disequilibrium gene flow genetic drift genetic variation genome genotype frequencies haplotypes Hardy-Weinberg expectations Hardy-Weinberg principle heritability heterozygosity heterozygous homozygotes homozygous identical by descent inbreeding coefficient increase individuals initial interaction island lethal linkage LOD score males microsatellite microsatellite loci molecular mutation rate natural populations natural selection neutral mutation neutral theory nucleotide observed occur offspring percent phenotypic polymorphism population genetics predicted probability of fixation problem proportion protein quantitative traits quency random mating random variable recessive recombination relative fitnesses reproductive sample Section self-fertilization single locus small populations species stable studies subpopulations substitution rates summarize Table tion tree variance viability zero zygotes